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Brodland for his input and support in the biomechanical modeling, D. del Alamo for molecular analysis of the neur IF65 allele, L. Couturier for technical help, and M. Labouesse for critical reading. This work was funded by an ANR grant (ANR12-BSV2-0010-01 to F.S.) and by MINECO grants (BFU2016-75101-P and RYC-2014-15559 to V, C. was also supported by the Centro de Excelencia Severo Ochoa Award to the Institute of Bioengineering of Catalonia . G.P.-M. was supported by MENRT and FRM fellowships ,